1. Department of Forest and Environmental System, College of Forest & Environmental Sciences, Kangwon National University, Chuncheon, Republic of Korea;
2. Ecosystem Research Division, National Park Research Institute, Wonju, Republic of Korea;
3. Department of Genetic Engineering, Sungkyunkwan University, Suwon, Republic of Korea;
4. Division of Forest Science, College of Forest & Environmental Sciences, Kangwon National University, Chuncheon, Republic of Korea