1. School of Biotechnology, Binzhou University, Binzhou, Shandong, P. R. China;
2. Shandong Provincial Key Laboratory of Eco-Environmental Science for Yellow River Delta, Binzhou University, Binzhou, Shandong, P. R. China;
3. Shandong Provincial Engineering and Technology Research Center for Wild Plant Resources Development and Application of Yellow River Delt, Binzhou University, Binzhou, Shandong, P. R. China;
4. BinZhou Municipal Bureau of Agriculture, Binzhou, Shandong, P. R. China