1. Red Sea Research Center, Division of Biological and Environmental Science and Engineering, King Abdullah University of Science and Technology, Thuwal, Saudi Arabia;
2. ARC Centre of Excellence for Coral Reef Studies, James Cook University, Townsville, QLD, Australia;
3. Department of Biotechnologies and Bioscience, University of Milano-Bicocca, Milan, Italy;
4. Hawaii Institute of Marine Biology, Kaneohe, HI, USA