1. Beijing Key Laboratory of Ornamental Plants Germplasm Innovation & Molecular Breeding, National Engineering Research Center for Floriculture and College of Landscape Architecture, Beijing Forestry University, Beijing, China;
2. Beijing Agro-Biotechnology Research Center, Beijing Academy of Agriculture and Forestry Sciences, Beijing, China;
3. Beijing Key Laboratory of Agricultural Genetic Resources and Biotechnology, Beijing, China;
4. Biological Sciences, University of California, Irvine, America