1. Department of Biological Science, Graduate School of Science, Hiroshima University, Hiroshima, Japan
2. Miyajima Natural Botanical Garden, Graduate School of Integrated Sciences for Life, Hiroshima University, Hiroshima, Japan
3. Program of Basic Biology, Graduate School of Integrated Sciences for Life, Hiroshima University, Hiroshima, Japan
4. Department of Botany, National Museum of Nature and Science, Ibaraki, Japan
5. Hattori Botanical Laboratory, Miyazaki, Japan