1. Department of Biological Sciences and Biotechnology, Botswana International University of Science and Technology, Palapye, Botswana;
2. Department of Ecology and Resource Management, University of Venda, Thohoyandou, South Africa;
3. Kansas Biological Survey and the Natural History Museum, Biodiversity Institute, Kansas University, Lawrence, Kansas, USA;
4. Centre for Ecological Genomics and Wildlife Conservation, Department of Zoology, University of Johannesburg, Auckland Park, South Africa