1. Zhanjiang City State Key Laboratory of Marine Ecology and Environment, Fisheries College, Guangdong Ocean University, Zhanjiang, China;
2. Animal Science College, South China Agricultural University, Guangzhou, China;
3. College of Life Science and Technology, Huizhou College, Huizhou, Guangdong, China;
4. College of Life Science and Technology, Hainan Tropical Ocean University, Sanya, Hainan, China