1. Leetown Science Center, Aquatic Ecology Branch, Natural Systems Analysts, Inc, Kearneysville, WV, USA;
2. United States Geological Survey, Virginia Water Science Center, Richmond, VA, USA;
3. Virginia Natural Heritage Program, Richmond, VA, USA;
4. United States Geological Survey, Leetown Science Center, Aquatic Ecology Branch, Kearneysville, WV, USA