1. Research Center for Marine Science, Hebei Normal University of Science and Technology, Qinhuangdao, China
2. Key Hebei Key Laboratory of Ocean Dynamics, Resources and Environments, Qinhuangdao, China
3. Key Laboratory of Tropical Marine Bio-resources and Ecology, Guangdong Provincial Key Laboratory of Applied Marine Biology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China