1. Division of Genetic Resources, Honam National Institute of Biological Resources, Mokpo, Republic of Korea
2. Division of Zoology, Honam National Institute of Biological Resources, Mokpo, Republic of Korea
3. Department of Applied Biology, College of Agriculture and Life Sciences, Chonnam National University, Gwangju, Republic of Korea
4. Scripps Institution of Oceanography, University of California San Diego, La Jolla, CA, USA