1. Department of Biodiversity and Evolutionary Biology, Museo Nacional de Ciencias Naturales (MNCN-CSIC), Madrid, Spain
2. A.N. Severtsov Institute of Ecology and Evolution Russian Academy of Sciences, Moscow, Russia
3. Laboratories of Analytical Biology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA
4. Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA