1. State Key Laboratory of Biocontrol, Institute of Aquatic Economic Animals and Guangdong Provincial Key Laboratory for Aquatic Economic Animals, School of Life Sciences, Sun Yat-Sen University, Guangzhou, China;
2. South China Sea Bio-Resource Exploitation and Utilization Collaborative Innovation Center, Guangzhou, China;
3. Marine Fisheries Development Center of Guangdong Province, Huizhou, China