1. Department of Biology, University of the Balearic Islands, Palma de Mallorca, Illes Balears, Spain;
2. IMEDEA (CSIC-UIB), Mediterranean Institute for Advanced Studies, Esporles, Illes Balears, Spain;
3. Universidad Nacional del Altiplano, Puno, Peru;
4. Department of Animal Ecology and Systematics, Justus Liebig University Giessen, Giessen, Germany