1. College of Marine Science and Biological Engineering, Qingdao University of Science and Technology, Qingdao, PR China
2. CAS Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, PR China
3. University of Chinese Academy of Sciences, Beijing, PR China;
4. Laboratory for Marine Ecology and Environmental Science, Qingdao National Laboratory for Marine Science and Technology, Qingdao, PR China
5. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, PR China