1. Fourth institute of Oceanography, Ministry of Natural Resources, Beihai, China
2. Institute of Marine Drugs, Guangxi University of Chinese Medicine, Nanning, China
3. Guangxi Engineering Technology Research Center for Marine Aquaculture, Guangxi Institute of Oceanology Co., Ltd, Beihai, China
4. Guangxi Key Laboratory of Aquatic Genetic Breeding and Healthy Aquaculture, Guangxi Academy of Fishery Sciences, Nanning, China