1. College of Life Sciences, Chongqing Normal University, Chongqing, China
2. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization & Ecological Restoration and Biodiversity Conservation Key Laboratory of Sichuan Province, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, China
3. College of Life Sciences, University of Chinese Academy of Sciences, Beijing, China