1. College of Ecological Environment and Resources, Qinghai Nationalities University, Xining, China
2. Qinghai Provincial Key Laboratory of Highvalue Utilization of Characteristic Economic Plants, Xining, China
3. College of Agriculture and Animal Husbandry, Qinghai University, Xining, China
4. Key Laboratory of Restoration Ecology in Cold Region of Qinghai Province, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining, China