1. Faculty of Agriculture, Forest and Food Engineering, Yibin University, Yibin, China
2. Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, China
3. Institute of Qinghai-Tibetan Plateau, Southwest Minzu University, Chengdu, China
4. University of Chinese Academy of Sciences, Beijing, China
5. Institute of Zoology, Chinese Academy of Sciences, Beijing, China