1. Department of Botany, National Museum of Nature and Science, Ibaraki, Japan
2. Hattori Botanical Laboratory, Miyazaki, Japan
3. Cooperative Research Fellow of Natural History Museum and Institute, Chiba, Japan
4. Center for Molecular Biodiversity Research, National Museum of Nature and Science, Ibaraki, Japan
5. Miyajima Natural Botanical Garden, Graduate School of Integrated Sciences for Life, Hiroshima University, Hiroshima, Japan