1. College of Forestry, Fujian Agriculture and Forestry University, Fuzhou, China;
2. Camellia Oleifera Research Center at College of Forestry, Fujian Agriculture and Forestry University, Fuzhou, China;
3. Key Laboratory of National Forestry and Grassland Administration for Orchid Conservation and Utilization at College of Forestry, Fujian Agriculture and Forestry University, Fuzhou, China;
4. College of Landscape, Fujian Agriculture and Forestry University, Fuzhou, China