1. Hainan Academy of Ocean and Fisheries Sciences, Haikou, People’s Republic of China;
2. Hainan Provincial Key Laboratory of technology for tropical seawater aquaculture, Haikou, People’s Republic of China;
3. Laboratory of Genetics and Breeding of Marine Organism, College of Marine Life Sciences, Ocean University of China, Qingdao, People’s Republic of China;
4. Marine Biology Institute, Shantou University, Shantou, People’s Republic of China