1. Key Laboratory for Forest Resources Conservation and Utilization in the Southwest Mountains of China, Ministry of Education, Southwest Forestry University, Kunming, Yunnan Province, China;
2. Wildlife Conservation Institute, Central South University of Forestry and Technology, Changsha, Hunan Province, China;
3. Key Laboratory for Conserving Wildlife with Small Populations in Yunnan, Southwest Forestry University, Kunming, Yunnan Province, China;
4. Faculty of Biodiversity Conservation, Southwest Forestry University, Kunming, Yunnan Province, China