1. Laboratory of Systematic and Evolutionary Botany and Biodiversity, College of Life Sciences, Zhejiang University, Hangzhou, Zhejiang, P. R. China;
2. Ministry of Education, Key Laboratory for Biodiversity Science and Ecological Engineering, Institute of Biodiversity Science, Fudan University, Shanghai, P. R. China;
3. Department of Environment and Forest Resources, Chungnam National University, Daejeon, South Korea