1. Department of Environmental Science, Faculty of Natural Resources & Desert Studies, University of Yazd, Yazd, Iran
2. Department of Biology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran
3. Research Department of Zoological Innovations (RDZE), Faculty of Sciences, Institute of Applied Zoology, Ferdowsi University of Mashhad, Mashhad, Iran