1. Afromontane Research Unit and Department of Geography, University of the Free State, South Africa
2. Centre for Biological Control, Department of Zoology and Entomology, Rhodes University, South Africa
3. Afromontane Research Unit and Department Zoology and Entomology, University of the Free State, South Africa
4. Centre for Invasion Biology, Department of Botany and Zoology, Stellenbosch University, South Africa
5. Institute of Botany, Czech Academy of Sciences, Czech Republic