1. School of Biological and Health Sciences, Technological University Dublin, City Campus, Dublin 8, Ireland
2. Radiation and Environmental Science Centre, FOCAS Research Institute, Camden Row, Dublin 8, Ireland
3. School of Physics & Clinical & Optometric Sciences, Technological University Dublin, City Campus, Dublin 8, Ireland
4. Department of Biology, Life Sciences building, McMaster University, 1280 Main Street West, Hamilton, Ontario, L8S 4K1, Canada