1. Quantitative Plant Ecology and Biodiversity Research Laboratory, Department of Biology, Ferdowsi University of Mashhad, Mashhad, Iran
2. Herbarium FUMH, Department of Botany, Research Center for Plant Sciences, Ferdowsi University of Mashhad, Mashhad, Iran
3. Department of Botany, Hungarian Natural History Museum, Budapest, Hungary
4. Institute for Soil Sciences, Centre for Agricultural Research, Budapest, Hungary