1. College of Geographic Sciences, Qinghai Normal University, Xining, Qinghai, China
2. Qinghai Province Key Laboratory of Physical Geography and Environmental Processes, Qinghai Normal University, Xining, Qinghai, China
3. School of Life Sciences, Arizona State University, Tempe, AZ, USA
4. Department of Ecosystem and Conservation Sciences, University of Montana, Missoula, Montana, USA
5. Washington Department of Fish and Wildlife, Olympia, Washington , USA