1. Department of Ecology, FSNV-STU, University of Guelma, Guelma, Algeria
2. Laboratoire de Conservation des Zones humides, University of Guelma, Guelma, Algeria
3. Department of Biology, University of Constantine, Constantine, Algeria
4. Department of Botany and Microbiology, King Saud University, Riyadh, Saudi Arabia
5. Department of Biology, University of Annaba, Annaba, Algeria