1. Department of Biology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran
2. Institute of Applied Zoology, Faculty of Science, Zoological Innovations Research Department, Ferdowsi University of Mashhad, Mashhad, Iran
3. Department of Biodiversity and Ecosystem Management, Environmental Sciences Research Institute, Shahid Beheshti University, G.C., Evin, Tehran, Iran