1. Caspian Institute of Biological Resources of the Dagestan Scientific Centre of the Russian Academy of Sciences, Makhachkala, Russia
2. Dagestan State University, Makhachkala, Russia
3. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, Moscow, Russia
4. Scientific Centre of Zoology and Hydroecology of the National Academy of Sciences of Armenia, Yerevan, Armenia