1. Department of Biology, School of Arts and Sciences, National University of Mongolia, Ulaanbaatar, Mongolia
2. Science Research Center, Hosei University, Fujimi, Tokyo, Japan
3. Graduate School of Science, Hokkaido University, Sapporo, Hokkaido, Japan
4. Center for Molecular Biodiversity Research, National Museum of Nature and Science,Tsukuba, Ibaraki, Japan
5. Museum Zoologicum Bogoriense, Zoology Division, Research Center for Biology, Indonesian Institute for Sciences, Bogor, Indonesia