1. Chongqing Key Laboratory of Economic Plant Biotechnology, College of Landscape Architecture and Life Science/Institute of Special Plants, Chongqing University of Arts and Sciences, Yongchuan, Chongqing, China
2. Yantai Lvyun Biotechnology Co., Ltd, Yantai, Shandong, China
3. College of Plant Protection, China Agricultural University, Beijing, China
4. Department of Biological Sciences, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, USA