1. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing, China
2. Laboratory for Marine Biology and Biotechnology, Pilot National Laboratory for Marine Science and Technology (Qingdao), Qingdao, China
3. CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, China
4. Deep Sea Research Center, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China