1. Biosciences College of life and Environmental Sciences, University of Exeter, Exeter, Devon, EX4 S4QD, UK
2. Pirbright Institute, Pirbright, Surrey, UK
3. Centre for Environment, Fisheries, and Aquaculture Science, Weymouth Laboratory, Weymouth, Dorset DT4 8UB UK
4. Istituto Zooprofilattico Sperimentale dell'Umbria e delle Marche, Laboratorio Nazionale di Riferimento Contaminazioni Batteriologiche dei Molluschi Bivalvi, Ancona, Italy
5. Department of Microbiology, Faculty of Science, Prince of Songkla University, Hat Yai 90110, Thailand