1. Shandong Key Laboratory of Animal Disease Control and Breeding, Institute of Animal Science and Veterinary Medicine, Shandong Academy of Agricultural Sciences, jinan, P. R. China
2. College of Life Sciences, Shandong Normal University, Jinan, China
3. Shandong Provincial Key Laboratory of Animal Cells and Developmental Biology, School of Life Science, Shandong University, Qingdao, China
4. College of Veterinary Medicine, Qingdao Agricultural University, Qingdao, China