1. North Carolina Cooperative Fish and Wildlife Research Unit, Department of Applied Ecology, North Carolina State University127 David Clark Labs, Campus Box 7617RaleighNorth Carolina27695‐7617USA
2. U.S. Geological Survey, North Carolina Cooperative Fish and Wildlife Research Unit, Department of Applied Ecology, North Carolina State University127 David Clark Labs, Campus Box 7617RaleighNorth Carolina27695‐7617USA