1. Department of Environmental Science, Rhodes University, Makhanda, South Africa
2. Centre for Invasion Biology, Department Environmental Science, Rhodes University, Makhanda, South Africa
3. Department of Conservation Ecology & Entomology, Stellenbosch University, Matieland, South Africa
4. Conservation Services, South African National Parks, Pretoria, South Africa
5. Centre for African Conservation Ecology, Nelson Mandela University, Gqeberha, South Africa