1. CIRAD, UMR TETIS, F-34398, Montpellier, France
2. TETIS, Univ Montpellier, AgroParisTech, CIRAD, CNRS, INRAE, Montpellier, France
3. CIRAD, UMR ASTRE, Montpellier, France
4. ASTRE, Univ Montpellier, CIRAD, INRAE, Montpellier, France
5. INRAE, UMR TETIS, Montpellier, France