1. Department of Forestry Genetics and Breeding, College of Forestry, Guizhou University, Guiyang, Guizhou, PR China
2. Department of Wildlife Conservation, Guizhou Botanical Garden, Guizhou Academy of Sciences, Guiyang, Guizhou, PR China
3. Department of Soil and Water Conservation, Institute of Mountain Resource, Guizhou Academy of Sciences, Guiyang, Guizhou, PR China
4. Key Laboratory for Biodiversity Conservation in Karst Mountain Area of Southwestern Chian of the National Forestry and Grass abd Administrations, Guiyang, Guizhou, PR China