1. Department of Biological Sciences, College of Natural and Computational Science, Mekdela Amba University, Tuluawliya, Ethiopia
2. Department of Industrial and Environmental Biotechnology, Institute of Biotechnology, University of Gondar, Gondar, Ethiopia
3. Department of Biological Sciences and Biotechnology, Faculty of Sciences, Botswana International University of Science and Technology, Palapye, Botswana
4. Bahir Dar Tannery Industry, Bahir Dar, Ethiopia