1. Faculty of Medicine, School of Clinical and Experimental Sciences, University of Southampton, Southampton, UK
2. Southampton NIHR Biomedical Research Centre, University Hospital Southampton NHS Foundation Trust and University of Southampton, Southampton, UK
3. Institute for Life Sciences, University of Southampton, Southampton, UK
4. GSK R&D, Stevenage, Hertfordshire, UK
5. Faculty of Medicine, School of Human Development and Health, University of Southampton, Southampton, UK
6. Biomedical Imaging Unit, Faculty of Medicine, University of Southampton, Southampton, UK