1. a Fisheries and Oceans Canada, Freshwater Institute , Winnipeg , R3T 2N6 , Canada
2. b State Key Laboratory of Tropical Oceanography, South China Sea Institute of Oceanology, Chinese Academy of Sciences , Guangzhou , 510301 , China
3. c Fisheries and Oceans Canada, Institut Maurice-Lamontagne, Mont-Joli , Québec , Canada , G5H 3Z4