1. Division of Psychiatry, The University of Edinburgh, Royal Edinburgh Hospital, Edinburgh, UK
2. Mrc Human Genetics Unit, Institute of Genetics and Cancer, the University of Edinburgh, UK
3. Centre for Genomic and Experimental Medicine, Institute of Genetics and Cancer, the University of Edinburgh, UK
4. Centre for Clinical Brain Sciences, The University of Edinburgh, UK
5. Aberdeen Biomedical Imaging Centre, The Institute of Medical Sciences, University of Aberdeen, UK
6. Imaging Science and Technology, School of Medicine, University of Dundee, Dundee UK
7. Centre for Cognitive Ageing and Cognitive Epidemiology, School of Philosophy, Psychology and Language Sciences, The University of Edinburgh, UK