1. Laboratory of Cellular Biochemistry, Department of Animal Resource Sciences/Veterinary Medical Sciences, The University of Tokyo, Tokyo, Japan
2. Synthetic Cellular Chemistry Laboratory, RIKEN, 2-1 Hirosawa, Wako, Saitama, Japan
3. Japan Science and Technology Agency ERATO Glycotrilogy Project, 2-1 Hirosawa, Wako, Saitama, Japan
4. Present address: Institute of Advanced Biosciences, Tokai University, 4-1-1 Kitakaname, Hiratsuka, Kanagawa, Japan
5. Waseda Research Institute for Science and Engineering, Waseda University, Tokyo, Japan