1. a ARC-Grain Crops Institute , Private Bag X1251, Potchefstroom , 2520 , Republic of South Africa
2. b Laboratory for Microscopy and Micro-Analysis , University of Pretoria , Pretoria , 0002 , Republic of South Africa
3. c Department of Plant Production and Soil Science , University of Pretoria , Pretoria , 0002 , Republic of South Africa