1. a Faculty of Bioresources , Mie University , Tsu , 514 , Japan
2. d Department of Applied Biological Chemistry, Division of Agriculture and Agricultural Life Sciences , The University of Tokyo , Bunkyo-ku, Tokyo , 113 , Japan
3. b Kirin Brewery Co. Ltd. , Toride Plant , Toride , 302 , Japan
4. c Faculty of Agriculture , Yamaguchi University , Yamaguchi , 753 , Japan