1. Graduate School of Bioagricultural Science, Nagoya University, Nagoya, Japan
2. Arctic Research Center, Hokkaido University, Sapporo, Japan
3. Graduate School of Environmental Studies, Hokkaido University, Sapporo, Japan
4. Institute for Biological Problems of Cryolithozone Siberian Branch, Russian Academy of Science, Yakutsk, Russia
5. Biogeochemistry Educational and Scientific Training Center, North-Eastern Federal University, Yakutsk, Russia