1. Research Unit: Biodiversity and Valorization of Arid Areas Bioressources (BVBAA), Faculty of Sciences, Gabes University, Gabes, Tunisia
2. Centre for Environmental Research and Studies (CERS), Jazan University, Jazan, Saudi Arabia
3. Laboratoire des Symbioses Tropicales et M´editerran´eennes, Cirad/IRD/Inra/Agro-M/UM2 Campus-international de Baillarguet, Montpellier Cedex 5, France