1. Agriculture, Forestry and Fisheries Research Council, Tokyo, Japan
2. Tohoku Agricultural Research Center, National Agricultural and Food Research Organization, Morioka, Japan
3. Central Agricultural Experiment Station, Hokkaido Research Organization, Nakashibetsu, Japan
4. Dairy Research Center, Hokkaido Research Organization, Naganuma, Japan
5. Central Agricultural Research Center, National Agricultural and Food Research Organization, Tsukuba, Japan